Hyperion (Mickey Rourke), the king of Crete, declares war on Olympus after the gods fail to answer his prayers to save his family from illness, and starts searching for the Epirus Bow, a powerful weapon created by Ares (Daniel Sharman), which he intends to use to release the Titans from Mount Tartarus and destroy the gods.
Hyperion attacks several holy places in search of the Epirus Bow, which was lost on Earth during the last war between the gods and the Titans, and after kidnapping a virgin oracle priestess called Phaedra (Freida Pinto) to help him, advances towards a nearby village, where the young Theseus (Henry Cavill) lives with his mother. Theseus' mother conceived him after being raped, and he is thus considered a bastard and prevented from evacuating the village with its other inhabitants by the soldier Lysander (Joseph Morgan). Theseus defeats him in combat, which he was taught by an old man (John Hurt), and Lysander is discharged by his commanding officer. Infuriated, he offers his services to Hyperion.
The old man is actually Zeus (Luke Evans) in disguise. He meets with Ares, Poseidon (Kellan Lutz), Athena (Isabel Lucas), Apollo (Corey Sevier) and Heracles (Steve Byers), and tells them that they can't directly interfere with the conflict until the Titans are released from Tartarus. Meanwhile, Theseus returns to the village only to discover that Hyperion and his army have already attacked. Hyperion murders Theseus' mother in front of him and sends him to work as a slave in the mines, where Theseus befriends another prisoner, the thief Stavros (Stephen Dorff).
Phaedra has a vision of Theseus standing as an ally to Hyperion after accidentally touching him and tells Stavros to make sure Theseus is always by his side. She and the other captured oracles later start a riot, in which she, Theseus, and Stavros escape alongside other slaves. Theseus decides to go after Hyperion and the others follow him. They attack a boat filled with Hyperion's men, but are overpowered. They are however rescued when Poseidon causes a tidal wave which knocks Hyperion's men out. On their way out, Phaedra has the same vision, noticing a shrouded body. She determines that this means Theseus needs to return to his village and bury his mother.
In the process, he finds the Epirus Bow, but is attacked by Hyperion's soldier, the Minotaur (Robert Maillet) while the others are attacked by soldiers. The Minotaur was warned of Theseus' presence by Lysander. Theseus finally defeats the Minotaur and rescues his allies using the Epirus Bow, before collapsing due to being poisoned by the Minotaur. Phaedra takes him back to his house, where she cures him and they make love, which causes her to lose her premonitory powers.
The group returns to Phaedra's temple, only to discover that it is filled with Hyperion's soldiers. They capture the group and steal the Epirus Bow, forcing Ares to intervene and save them. Athena comes from Olympus as well. Zeus sees this as treachery, and is ready to kill Athena, but he spares her life. Ares, however, does not back down, and says that he had to do what he did. An infuriated Zeus uses a fire chain to knock Ares into the marble wall killing him. Athena is able to provide horses for the group (under her father's ultimatum that he'd do to her what he did to Ares if she ever aids the group again) which they use to ride to Mount Tartarus which is under attack by Hyperion's forces. Hyperion releases the Titans using the Epirus Bow, and they are confronted by the gods, while Theseus and his allies join forces with the Athenian army present.
Theseus enters the temple and fights Hyperion. He is stabbed multiple times, but is able to kill Hyperion by stabbing him in the neck before dying. Meanwhile, the gods are overpowered by the Titans, who kill Stavros, Apollo, Athena , Heracles and Poseidon. Before Zeus returns to Mount Olympus with his dead daughter, he collapses the temple upon the Titans, killing them as well as most of Hyperion's forces, while the rest are crushed by the collapsing temple. Two beams are seen ascending to Olympus; Zeus carrying Athena. Theseus also ascends to the skies as a beam.
Theseus is welcomed into Mount Olympus by the gods for his sacrifice, and is awarded with a son from his night with Phaedra. The boy inherits his mother's premonitions and is told by the old man not to fear them, though his visions show that in the future, there will be a war between the gods and the Titans, with Theseus leading the gods in battle on Mount Olympus. The old man tells him that his time for war and glory will come.
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